Archaegenetics in Croatia and origin of Croats

This article will include a collection and analysis of archaeogenetic samples from Croatia. The first version will be published soon and is under construction until the removal of this notice. For now, please check the "Ancient and medieval archaeogenetics and origin of Croats in the case of Velim, Croatia" (PDF file at Academia.edu), which is an analysis of the ancient and medieval archaeogenetics of the archaeological site of Velim in Croatia and the related origin of the Croats (per Gretzinger et al. 2025 and previously published scientific papers), and "Ancient DNA database from Croatia" (Excel file at Academia.edu).

First will be an abstract of observed data (Y-DNA, mtDNA, and atDNA), and sections dealing with the medieval ancestry and its relation to the modern Croats, and at the end, the list of all archaeogenetic samples with related data (ID, location, time period, mtDNA, and Y-DNA haplogroup). The current ancient DNA database contains 465 samples. Of these, around 200 have Y-DNA haplogroup assignments and 395 have mtDNA haplogroup assignments. There may be sample bias in certain periods, for example, the Neolithic is strongly represented by the Beli Manastir–Popova Zemlja site, the Chalcolithic by the Potočani site, the Bronze Age by Bezdanjača Cave and sites in the Cetina Valley, and the Roman period by Radošević (Hvar) and Bubi’s Cave (BBC). However, the Middle Ages are represented by more diverse sites like Velim (VEM).

Y-DNA haplogroups:

There's no Y-DNA sample for the Mesolithic Age, but considering the Iron Gates Mesolithic archaeological culture from eastern Serbia and western Romania, it could be considered that the predominantly present haplogroups were I2a-L460, C-F3393, and R1b1a-L754, which are typical of Western or Eastern European hunter-gatherers.

In the Neolithic Age, Anatolian Neolithic farmers arrived, assimilated previous populations, and were predominantly represented by haplogroup G2a (43%) with some C (25%) and I2 (18%), and a few early E1b1b and J2a. The predominance of G2a (68%) and the top three haplogroup combinations persisted in the Neolithic and Chalcolithic Age, and the mentioned haplogroup I2 was represented only by I2-M223, I2-L596, and I2-P37.2>CTS595>L1286 clades, but the I2 almost vanished in subsequent ages.

In the Bronze Age, a large-scale migration and conquest of Europe by the Indo-Europeans, seen with predominance of R1b (33%) and J2b (47%), and diminishing of a still significant minority of G2a (16%). In the Iron Age became even more predominant (66%) haplogroup J2b(-L283) specifically related to the Illyrians. With their downfall and conquest by the Romans, the population's paternal genetic heritage became very heterogeneous, with J2b diminishing to 22% and the rise of J2a (17%), E1b1b (15%), and the appearances of barbarian I1, R1a(-Z93), and N. In the early medieval period, the genetic landscape changed drastically once again, with the arrival of the early Slavs. Most prominent haplogroups became I2-M423(>CTS10228>Y3120) and R1a, both with around 20%, which persisted and/or rose until the modern age (36% and 23%), while Illyrian J2b(>L283) almost vanished (~2%). For the modern population, a frequency was used from a sample population from Purps et al. 2014 with a similar sample size as the current ancient sample size.

Ancient DNA Y-DNA haplogroup frequencies

mtDNA haplogroups:

mtDNA haplogroups, due to their slow mutation rate and widespread distribution, are difficult to analyze in the context of historical migrations, especially of small-distance and time periods. For detailed analysis and conclusions, phylogenetically younger subclades are needed. As there's a lack of mtDNA samples for the Mesolithic Age as well (only one U5), considering the Iron Gates Mesolithic samples, probably were also present U4, K1, and H(13) haplogroups. In comparison to modern age haplogroups' frequencies – based on superficial macro-haplogroup phylogeny - it could be observed, although only presumably, that the modern mtDNA haplogroup composition was partly formed in the Neolithic Age and most prominently in the Iron Age/Roman Age (at least in the predominance of haplogroup H). However, some studies like Davidovic et al. 2014/2017/2020 about Serbian mtDNA haplogroup frequencies claim the existence of pre-Slavic and Slavic gene pool lineages, further research is needed. In the case of early medieval Velim from Gretzinger et al. 2025, compared to pre-Slavic samples from Croatia, showed ancient continuity of only around 30%, probably indicating a medieval ancestry of most mtDNA haplogroups in modern populations. In the statistics of Croatia, there can still be seen some trends, diminishing of haplogroup T since the Bronze Age or Roman Age and Modern Age, of haplogroup K since the Bronze Age, higher frequency of haplogroup R in Modern Age (if Barbaric et al. 2020 made the right correction), and higher diversity of haplogroup U subclades in Modern Age. According to Šarac et al. 2014, based on frequencies, the Croatian sub-populations of South, East, Bosnia and Herzegovina plot together while West and North (or North only) plot together near Slovenia (and all Croatian sub-populations somewhere between Slovakia, Czechia, Hungary, Poland, Ukraine and Germany), while according to Barbaric et al. 2020 the Croatian population plots between Slovakia, Hungary and Romania.

Ancient DNA mtDNA haplogroup frequencies

Based on matching and checking YFull and FTDNA MTree with criteria, whether the haplogroup subclade has major prevalence in modern Eastern-Central Europe and significant presence in Southern Europe (partly excluding South Slavic countries, but still taking into consideration):

BBC = 34 samples

VEM = 61 samples (with high diversity of subclades)

BBC predominantly has mtDNA haplogroups which were previously confirmed in Illyrian samples from the Bronze and Iron Age (HV0e, H1, H5, H6a, H7, K1a3, J1c), but also which didn't (e.g. 5x I5c, which is seemingly Central European and all those BBC samples have Celtic Y-DNA haplogroup G-L497, or 4x H33c).

There's a possible match between BBC and early medieval VEM only at:

2x H5 vs 5x H5a (H5a, H5a1, H5a1m, H5a2, H5a2f)

J1c2 vs J1c, J1c2k, J1c2u1a

W1i1a/W1r vs W1e1a1

HV0e vs HV0e

= 10/61 (16.3%)

When we add 131 mtDNA sample results from Bronze, Iron, and Roman age Croatia to the BBC, expanding the number to 165 samples, then the number of possible matches could rise a bit (with some doubts with a question mark):

2x H5 vs 5x H5a (H5a, H5a1, H5a1m, H5a2, H5a2f)

J1c2 vs J1c, J1c2k, J1c2u1a

W1i1a/W1r vs W1e1a1

HV0e vs HV0e

J1c3/J1c3-a1* vs J1c3-a1

K1a4 Vs 2x K1a4c (K1a4c1, K1a4c1f)

U5/U5a (U5a1-a)

R0a (R0a1b) – West Asia (Semitic)

= 15/61 (24.5%)

Doubtful:

3x H6a1a (H6a1a12, H6a1a4b3; ?) – Eastern Europe

2x H3-c (?) – Southern Europe

U5a1b1c (?) – Eastern Europe

H1a (?) – Northern-Western Europe

H84a (?) – Southern Europe

V6 (?) – West Asia (Semitic)

U8b (?) – too old/uncertain

= 19/61 (31.1%)

The remaining subclades without any ancient match and thus possibly brought during the migration period are:

3x H6a1a (H6a1a12, H6a1a4b3) – Eastern Europe

U5a1b1c – Eastern Europe

T2a1a1b – Eastern Europe

H13a1c – Eastern Europe, West & Central Asia

2x K1-a – too old/uncertain

3x T2b5 – Western & Central & Eastern Europe

V7a1 – Northern Europe

H32 – Southern & Western Europe

U7b19 – Central & Eastern Europe

R13 – too old/uncertain / Southern Europe

2x V4 – Western & Central & Northern Europe

U5b although exist before, not these subclades:

(U5b1l – Central Europe

2x U5b1e1 – Eastern Europe

U5b1-a1 – Southern & Eastern Europe

2x U5b1b1a – Eastern & Northern Europe)

2x I5a (I5a1) – too old/uncertain / all over Europe

N14 – uncertain

H36a – Eastern & Central Europe

H1f-a – Northern Europe

H-d17 – Southern Europe

M31a2b2 - India

U4d1a1a1 – Northern Europe

B5a1c1a4 – East Asia (China, Vietnam)

I3a – Eastern & Southern Europe

I14 – Southern Europe & North Africa

U2e1b1a – Eastern Europe

2x H1c (H1c22) – Eastern & Northern & Central & Western Europe

HV2a1a – West Asia & over all Europe

K1a2 – over all Europe

U4c – Eastern & Southern Europe

H2a2a1i – Northern & Eastern Europe

J1b1a1b – Eastern & Northern & Central Europe

Such analysis and comparison of mtDNA haplogroups with good enough phylogenetic resolution show a continuity of only up to 30% and that the vast majority of mtDNA haplogroups not found in the pre-Slavic period are not found today in Southern Europe as well, but can be found in Eastern Europe, proving, alongside atDNA admixture & IBD and Y-DNA haplogroups, that there was a large movement of Slavic people without gender bias.

When comparing the mtDNA subclades from Velim (using the listed assignments in the paper as is easier to compare, although there's a slight difference/better in assignments posted at the Genarchivist forum), with EPolish, WUkrainian, EGermany Slavic sites, there's a match on:

H1a in EGermany (NDW, OBM, SDN)

H1c in EGermany (NDW, OBM, SDN) and Poland (GRK)

H36 in EGermany (NDW) and Ukraine (PDH)

H5a2 in EGermany (OBM)

H6a1a level in EGermany (NDW, OBM, SDN) and Ukraine (KRW)

J1b1a1 level in Poland (GRK) and Ukraine (GRK)

J1c5 in EGermany (NDW)

K1a2 in Ukraine (KRW)

K1a4c1 in EGermany (SDN)

T2b4 level in EGermany (NDW) and Ukraine (PDH)

U2e1b1 in Latvia SP (LAU), while on level U2e1 is Ukraine (KRW)

U5a1b1c level in Ukraine (KRW)

U5b1b1a in EGermany (OBM)

U5b1e1 in EGermany (NDW, SDN)

V7a level in EGermany (NDW)

W1e1a in EGermany (SDN)

The point isn't finding matches to see the percentage of non-shared maternal lineages (it is substantial between all of them), but simply finding matches and whether they have any significance in a similar fashion to a lack of shared paternal Y-DNA haplogroup lineages between these Slavic sites. In contrast to the comparison of the pre-Slavic and Slavic period mtDNA haplogroups in Croatia (as periods differentiate the populations genetically), and an additional share of maternal lineages between non-Velim sites (e.g. H6a1a4 OBM-KRW, level of J1c2e KRW-SDN, level of J2a1a1a NDW-KRW, T2b4 NDW-PDH, U3b NDW-GRK, V1a1b NDW-KRW, W6a PDH-NDW), the same lineages are shared in multiple sites, showing to merely be evidence of the common source population and tribal connections predominantly by females in a strongly patrilineal tribal early Slavic world.

atDNA admixture and IBD:

PCA and Admixture of the Copper Age Potočani individuals with PCA of other Neolithic & Bronze & Iron Age samples compared to present-day Croats, per Novak, Olalde, Ringbauer et al. 2021

West Eurasian PCA's, Admixture and genetic affinities of Roman-age Mursa, Croatian Iron Ages and modern Croatian genomes, per Novak et al. 2025

The genetic legacy of the Slavic expansion (black, in Croatia roughly 70%), per Gretzinger et al. 2025

Early Slavic admixture based on Velim samples (G25 profile), per ph2ter

Velim samples IBD > 20 cM with a EUkrainian, Polish, and EGermany cluster of Slavic sites; nevertheless, if taken into consideration IBD > 20 or 12 cM and sample dating (pre and post 900/1000 AD), and the fact that the sum is influenced by sample number/bias, the averages show significantly stronger and deeper connection with Ukrainian and Polish rather than with the German cluster

Based on the "Extended Data Fig. 10: Changes in genetic relatedness between and within populations", the IBD network indicates that the EPolish & WUkrainian samples make a wide cluster which encompasses all other clusters (EGermany, Czechia, Croatia), each encircled by another smaller EPolish & WUkrainian cluster, proving that the early Slavic migration had a common source in Eastern Europe, vector source of the Croatian cluster was not in Czechia, but in Eastern Europe (EPoland & WUkraine), but further research with more details will proves or disapprove this preliminary conclusions

List:

1 I1875, Szécsényi-Nagy et al. 2015, Dalmatia, Vela Spila, Mesolithic, 7308-7027 BCE, U5b2a5

2 VINK 1, Szécsényi-Nagy et al. 2015, Continental, Vinkovci, Early Neolithic, Starčevo culture, K1a

3 VINK 2, Szécsényi-Nagy et al. 2015, Continental, Vinkovci, Early Neolithic, Starčevo culture, J1c

4 VINK 3, Szécsényi-Nagy et al. 2015, Continental, Vinkovci, Early Neolithic, Starčevo culture, HV0

5 VINK 4, Szécsényi-Nagy et al. 2015, Continental, Vinkovci, Early Neolithic, Starčevo culture, n/a, J?

6 VINK 5, Szécsényi-Nagy et al. 2015, Continental, Vinkovci, Early Neolithic, Starčevo culture, K1a, G2a

7 VINK 6, Szécsényi-Nagy et al. 2015, Continental, Vinkovci, Early Neolithic, Starčevo culture, n/a, T? 

8 VINJ 1, Szécsényi-Nagy et al. 2015, Continental, Vinkovci Jugobanka, Early Neolithic, Starčevo culture, T2b 

9 VINJ 2, Szécsényi-Nagy et al. 2015, Continental, Vinkovci Jugobanka, Early Neolithic, Starčevo culture, K 

10 VINJ 3, Szécsényi-Nagy et al. 2015, Continental, Vinkovci Jugobanka, Early Neolithic, Starčevo culture, V 

11 VINJ 4, Szécsényi-Nagy et al. 2015, Continental, Vinkovci Jugobanka, Early Neolithic, Starčevo culture, V6, I2a1-P-37.2

12 VUKG 1, Szécsényi-Nagy et al. 2015, Continental, Vukovar Gimnazija, Early Neolithic, Starčevo culture, J1c, G2a

13 VUKG 2, Szécsényi-Nagy et al. 2015, Continental, Vukovar Gimnazija, Early Neolithic, Starčevo culture, n/a, n/a 

14 VUKG 3, Szécsényi-Nagy et al. 2015, Continental, Vukovar Gimnazija, Early Neolithic, Starčevo culture, J1c 

15 VUKG 4, Szécsényi-Nagy et al. 2015, Continental, Vukovar Gimnazija, Early Neolithic, Starčevo culture, T2b 

16 I3498, Mathieson et al. 2018, Continental, Beli Manastir-Popova zemlja, Early Neolithic, 5837-5659 calBCE, Starčevo culture, U8b1b1, C

17 I3433, Mathieson et al. 2018, Dalmatia, Zemunica Cave, Early Neolithic, 5976-5751 BCE, Croatia_Cardial_Neolithic, H1 

18 I3947, Mathieson et al. 2018, Dalmatia, Zemunica Cave, Early Neolithic, 5986-5786 BCE, Croatia_Cardial_Neolithic, K1b1a, C1a2-V20

19 I3948, Mathieson et al. 2018, Dalmatia, Zemunica Cave, Early Neolithic, 6005-5814 BCE, Croatia_Cardial_Neolithic, N1a1, E1b1b1a1b1

20 I5071, Mathieson et al. 2018, Istria, Kargadur, Early Neolithic, 5670-5571 BCE, Croatia_Impressa_EN, H5a 

21 I5072, Mathieson et al. 2018, Istria, Kargadur, Early Neolithic, 5641-5560 BCE, Croatia_Impressa_EN, H7c, G2a2a1

22 I26738, Patterson et al. 2022, Continental, Gornja Vrba-Savsko polje, Middle Neolithic, 5450-4900 BCE, Croatia_MN, T2b/T2b81b 

23 I5077, Mathieson et al. 2018, Continental, Osijek, Middle Neolithic, 5207-4945 BCE, Sopot culture U5a1a2/U5a1c1e, G2a2a1

24 I5078, Mathieson et al. 2018, Continental, Osijek, Middle Neolithic, 4692-4546 BCE, Sopot culture, H10/H10-a*, J2a1

25 I7399, Lazaridis et al. 2022, Continental, Vinkovci-Vinka, Middle Neolithic, 5216-5045 BCE, HRV_Vinkovci_MN, V1 

26 I7398, Lazaridis et al. 2022, Continental, Vinkovci-Vinka, Middle Neolithic, 5100-4750 BCE, HRV_Vinkovci_MN, H84, C1a2-V86

27 I7538, Lazaridis et al. 2022, Continental, Vinkovci-Vinka, Middle Neolithic, 5100-4750 BCE, HRV_Vinkovci_MN, V1 

28 I4167, Mathieson et al. 2018, Continental, Beli Manastir-Popova zemlja, Middle-Late Neolithic, 4790-4558 BCE, Sopot culture, U5b2b, I

29 I4168, Mathieson et al. 2018, Continental, Beli Manastir-Popova zemlja, Middle-Late Neolithic, 4763-4536 BCE, Sopot culture, N1a1 

30 I26741, Patterson et al. 2022, Continental, Osijek-Hermanov vinograd, Middle Neolithic, 4713-4548 BCE, Sopot culture, T2b 

31 POP02, Freilich et al. 2021, Continental, Beli Manastir-Popova zemlja, Middle Neolithic, 4700-4300 BCE, Sopot culture, K1a/K1-a4x 

32 POP04, Freilich et al. 2021, Continental, Beli Manastir-Popova zemlja, Middle Neolithic, 4700-4300 BCE, Sopot culture, H, J-M304 

33 POP05, Freilich et al. 2021, Continental, Beli Manastir-Popova zemlja, Middle Neolithic, 4700-4300 BCE, Sopot culture, K1a5 

34 POP06, Freilich et al. 2021, Continental, Beli Manastir-Popova zemlja, Middle Neolithic, 4700-4300 BCE, Sopot culture, K2b1/K2b1c 

35 POP07/I4167, Freilich et al. 2021, Continental, Beli Manastir-Popova zemlja, Middle Neolithic, 4790-4558 BCE, Sopot culture, U5b2b, I2a2a-M223

36 POP08, Freilich et al. 2021, Continental, Beli Manastir-Popova zemlja, Middle Neolithic, 4700-4300 BCE, Sopot culture, U8b1a1/U8b1a1a3b* 

37 POP09, Freilich et al. 2021, Continental, Beli Manastir-Popova zemlja, Middle Neolithic, 4700-4300 BCE, Sopot culture, K1a4 

38 POP11, Freilich et al. 2021, Continental, Beli Manastir-Popova zemlja, Middle Neolithic, 4700-4300 BCE, Sopot culture, T2b3/T2b11b 

39 POP12, Freilich et al. 2021, Continental, Beli Manastir-Popova zemlja, Middle Neolithic, 4700-4300 BCE, Sopot culture, T2b/T2b55a1* 

40 POP13, Freilich et al. 2021, Continental, Beli Manastir-Popova zemlja, Middle Neolithic, 4700-4300 BCE, Sopot culture, T2c1d1 

41 POP14/I4168, Freilich et al. 2021, Continental, Beli Manastir-Popova zemlja, Middle Neolithic, 4763-4536 BCE, Sopot culture, N1a1a1 

42 POP16, Freilich et al. 2021, Continental, Beli Manastir-Popova zemlja, Middle Neolithic, 4700-4300 BCE, Sopot culture, N1a1a1a2 

43 POP19, Freilich et al. 2021, Continental, Beli Manastir-Popova zemlja, Middle Neolithic, 4700-4300 BCE, Sopot culture, N1a1a1a3/N1a1a1a3b 

44 POP24, Freilich et al. 2021, Continental, Beli Manastir-Popova zemlja, Middle Neolithic, 4700-4300 BCE, Sopot culture, K1a1a, I2a2a-M223

45 POP27, Freilich et al. 2021, Continental, Beli Manastir-Popova zemlja, Middle Neolithic, 4700-4300 BCE, Sopot culture, T2b21 

46 POP30, Freilich et al. 2021, Continental, Beli Manastir-Popova zemlja, Middle Neolithic, 4700-4300 BCE, Sopot culture, T2b11, G2a2a-PF3147

47 POP33, Freilich et al. 2021, Continental, Beli Manastir-Popova zemlja, Middle Neolithic, 4603-4071 BCE, Sopot culture, K1a1, G2a2b2a1a1-PF3345

48 POP35, Freilich et al. 2021, Continental, Beli Manastir-Popova zemlja, Middle Neolithic, 4584-4458 BCE, Sopot culture, J2b1a5, C1a2b-Z38888

49 POP36, Freilich et al. 2021, Continental, Beli Manastir-Popova zemlja, Middle Neolithic, 4700-4300 BCE, Sopot culture, H, G2a2a-PF3147

50 I18827, Lazaridis et al. 2022, Continental, Gornja Vrba - Savsko polje, Middle Neolithic, 5000-4000 BCE, n/a, IJ 

51 I10045, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, H13b1+200, G2a2b2a1 

52 I10046, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, N1a1a1a3, G2a2b2a1 

53 I10047, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, U2, G2a2a1a 

54 I10048, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, U5b2c , G2a2b2a1a1c 

55 I10049, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, U2, G2a2b2a1 

56 I10050, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, T2b 

57 I10052, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, U5b1d1 

58 I10053, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, H5b, G2a2a1a2 

59 I10054, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, H, G2a2b2a1a1c 

60 I10055, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, K1a1 

61 I10056, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, K1a3a 

62 I10057, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, H7 

63 I10058, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, K1b1b1, G2a2b2a1a 

64 I10061, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, H5, G2a2b2a1 

65 I10062, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, HV, I2a1a(xI2a1a2) 

66 I10063, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, H, C1a2(xC1a2a1) 

67 I10064, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, J1 

68 I10065, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, H42 

69 I10067, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, H4a1 

70 I10068, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, T1a2, G2a2b2a1a1a 

71 I10069, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, T2f 

72 I10070, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, K1a3a 

73 I10071, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, H26 

74 I10072, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, N1a1a1, I2a1a(xI2a1a2) 

75 I10074, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, K1a3a 

76 I10271, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, T2b23, I2a2 

77 I10272, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, T2b 

78 I10273, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, N1a1a1a2 

79 I10274, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, T2b, C(xC1a1a1a,C1b3,C2) 

80 I10275, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, T2b 

81 I10291, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, T2b23 

82 I10292, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, n/a, n/a 

83 I10293, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, H4a1 

84 I10294, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, H, G2a2b2a1a1c

85 I10295, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, T2b, n/a 

86 I10297, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, X2b 

87 I10299, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, K1a4a1

88 I10303, Novak et al. 2021, Continental, Potočani, Middle Chalcolithic, 4200 BCE, Lasinja culture, N1a1a1

89 I20502, Lazaridis et al. 2022, Continental, Dakovo - Franjevac, Chalcolithic, 5000-3000 BCE, HRV_ChL, H5a, G2a2b2b1a1a2-Z7016

90 I5079, Mathieson et al. 2018/Patterson et al. 2022, Continental, Radovanci, Chalcolithic, 3710-3360 BCE, Croatia_Eneolithic/Croatia_N_Starcevo, J1c2/J1c2aa 

91 I3499, Mathieson et al. 2018, Continental, Beli Manastir-Popova zemlja, Chalcolithic, 2884-2666 BCE, Vučedol culture, T2e, R1b1a1a2a2/R-M12149/R1b1a1a2a2:Z2105

92 POP39, Freilich et al. 2021, Continental, Beli Manastir-Popova zemlja, Chalcolithic, 2859-2502 BCE, Croatia_Pop_CA, HV9/HV-b 

93 I2792, Mathieson et al. 2018/Patterson et al. 2022, Continental, Vucedol Tell, Late Chalcolithic-Early Bronze Age, 2872-2582 BCE, Vučedol culture, T2c1d1, G2a-Z6484

94 I4175, Mathieson et al. 2018/Patterson et al. 2022, Continental, Vucedol Tell, Late Chalcolithic-Early Bronze Age, 3000-2700 BCE, Vučedol culture, U4a 

95 JAG06, Freilich et al. 2021, Continental, Jagodnjak-Krcevine, Middle Bronze Age, 1800-1600 BCE, Croatia_Jag_MBA, T2b11, G2a2a1a2a2a1-Z31430/Y140827

96 JAG34, Freilich et al. 2021, Continental, Jagodnjak-Krcevine, Middle Bronze Age, 1879-1642 BCE, Croatia_Jag_MBA, K2a, G2a2a1a2a2a1-Z31430/Y140827

97 JAG58, Freilich et al. 2021, Continental, Jagodnjak-Krcevine, Middle Bronze Age, 1800-1600 BCE, Croatia_Jag_MBA, T2b11, G2a2a1a2a2a1-Z31430/Y140827

98 JAG78, Freilich et al. 2021, Continental, Jagodnjak-Krcevine, Middle Bronze Age, 1800-1600 BCE, Croatia_Jag_MBA, U5b1b1a/U5b1b1-b, G2a2a1a2a2a1-Z31430/Y140827

99 JAG82, Freilich et al. 2021, Continental, Jagodnjak-Krcevine, Middle Bronze Age, 1800-1600 BCE, Croatia_Jag_MBA, U2e1a1, G2a2a1-PF3148

100 JAG85, Freilich et al. 2021, Continental, Jagodnjak-Krcevine, Middle Bronze Age, 1800-1600 BCE, Croatia_Jag_MBA, K1b1b1 

101 JAG93, Freilich et al. 2021, Continental, Jagodnjak-Krcevine, Middle Bronze Age, 1800-1600 BCE, Croatia_Jag_MBA, U5a1g/U5a1-a* 

102 I11843, Lazaridis et al. 2022, Dalmatia, Cetina Valley,  Middle Bronze Age, 2000-1600 BCE, Cetina culture, T1a1, J2b2a1-Z597

103 I19017, Lazaridis et al. 2022, Dalmatia, Cetina Valley, Middle Bronze Age, 2000-1600 BCE, Cetina culture, H6a1a 

104 I18752, Lazaridis et al. 2022, Dalmatia, Cetina Valley, Middle Bronze Age, 2000-1600 BCE, Cetina culture, H13a2a, R1b1a1b1a-Z2118

105 I18088, Lazaridis et al. 2022, Dalmatia, Cetina Valley, Middle Bronze Age, 2000-1600 BCE, Cetina culture, H5, J2b2a-M241

106 I18747, Lazaridis et al. 2022, Dalmatia, Cetina Valley, Middle Bronze Age, 2000-1600 BCE, Cetina culture, U5a1a, J2b2a1a1a1b-Y15058

107 I19032, Lazaridis et al. 2022, Dalmatia, Cetina Valley, Middle Bronze Age, 2000-1600 BCE, Cetina culture, J1c1, J2b2a1a1a1b-Y15058

108 I19031, Lazaridis et al. 2022, Dalmatia, Cetina Valley, Middle Bronze Age, 2000-1600 BCE, Cetina culture, J1c1, J2-M102

109 I19030, Lazaridis et al. 2022, Dalmatia, Cetina Valley, Middle Bronze Age, 2000-1600 BCE, Cetina culture, J1c1

110 I19029, Lazaridis et al. 2022, Dalmatia, Cetina Valley, Middle Bronze Age, 2000-1600 BCE, Cetina culture, J1c1, J2b2a1-Z600

111 I19027, Lazaridis et al. 2022, Dalmatia, Cetina Valley, Middle Bronze Age, 2000-1600 BCE, Cetina culture, H 

112 I19026, Lazaridis et al. 2022, Dalmatia, Cetina Valley, Middle Bronze Age, 2000-1600 BCE, Cetina culture, J1c1, J2-M102

113 I19025, Lazaridis et al. 2022, Dalmatia, Cetina Valley, Middle Bronze Age, 2000-1600 BCE, Cetina culture, J1c1, J2b2a1-L283

114 I19022, Lazaridis et al. 2022, Dalmatia, Cetina Valley, Middle Bronze Age, 2000-1600 BCE, Cetina culture, H 

115 I19020, Lazaridis et al. 2022, Dalmatia, Cetina Valley, Middle Bronze Age, 2000-1600 BCE, Cetina culture, H6a1a 

116 I19019, Lazaridis et al. 2022, Dalmatia, Cetina Valley, Middle Bronze Age, 2000-1600 BCE, Cetina culture, H6a1a 

117 I19016, Lazaridis et al. 2022, Dalmatia, Cetina Valley, Middle Bronze Age, 2000-1600 BCE, Cetina culture, H6a1 

118 I18746, Lazaridis et al. 2022, Dalmatia, Cetina Valley, Middle Bronze Age, 2000-1600 BCE, Cetina culture, N1a1a1, J2b2a1a1a1b2-PH1602

119 I18745, Lazaridis et al. 2022, Dalmatia, Cetina Valley, Middle Bronze Age, 2000-1600 BCE, Cetina culture, HV0e 

120 I18748, Lazaridis et al. 2022, Dalmatia, Bogomolje, Middle Bronze Age, 1900-1600 BCE, HRV_BA, X2k, J2b2a1a1a1b-Y15058

121 I5080, Lazaridis et al. 2022, Dalmatia, Zavojane-Ravca, Middle Bronze Age, 1743-1623 BCE, HRV_BA, K1a4, J2b2a1a1a1b-Z38240

122 I18712, Lazaridis et al. 2022, Dalmatia, D. Ostrvica-Pasičine, Middle Bronze Age, 2500-800 BCE, HRV_BA, H1u, J2b2a1a1a1b2-PH1602

123 I5073, Lazaridis et al. 2022, Dalmatia, Koprivno, Middle Bronze Age, 1732-1542 BCE, HRV_BA, J2a1, J2b2a1a1a1b2-PH1602

124 I4331, Mathieson et al. 2018/Patterson et al. 2022, Dalmatia, Veliki Vanik, Early-Middle Bronze Age, 1631-1521 BCE, Croatia_EMBA, I1a1, J2b2a-L283

125 I4332, Mathieson et al. 2018/Patterson et al. 2022, Dalmatia, Veliki Vanik, Early-Middle Bronze Age, 1618-1513 BCE, Croatia_EMBA, W3a1 

126 I26726, Patterson et al. 2022, Dalmatia, Gudnja cave, Middle Bronze Age, 1507-1415 BCE, Croatia_MBA, J1c3-a1*, J2b2a1a1a1a1a -Z1297

127 I24342, Patterson et al. 2022, Dalmatia, Velim-Kosa, Middle Bronze Age, 1501-1421 BCE, Croatia_MBA, T2b23-a*, R1b1a1b1a1a2b1-L2

128 I26773, Patterson et al. 2022, Dalmatia, Mušego/Mon Sego, Middle Bronze Age, 1550-1200 BCE, Croatia_MBA, J1c2 

129 I26774, Patterson et al. 2022, Dalmatia, Mušego/Mon Sego, Middle Bronze Age, 1550-1200 BCE, Croatia_MBA, J1c2/J1c2ao 

130 I26893, Patterson et al. 2022, Dalmatia, Mušego/Mon Sego, Middle Bronze Age, 1550-1200 BCE, Croatia_MBA, U8b1b1d 

131 I18073, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 2000-1000 BCE, HRV_Bezdanjača_BA, U5b1c1 

132 I18082, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 2000-1000 BCE, HRV_Bezdanjača_BA, U5a1 

133 I18074, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 2000-1000 BCE, HRV_Bezdanjača_BA, H 

134 I18072, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 2000-1000 BCE, HRV_Bezdanjača_BA, H5n, R1b1a1b1a1a2b1c2-Y14088

135 I18087, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 2000-1000 BCE, HRV_Bezdanjača_BA, K1a 

136 I18078, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 2000-1000 BCE, HRV_Bezdanjača_BA, H5r 

137 I18079, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 2000-1000 BCE, HRV_Bezdanjača_BA, H46 

138 I18077, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 2000-1000 BCE, HRV_Bezdanjača_BA, U5a1a1 

139 I18085, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 2000-1000 BCE, HRV_MLBA, n/a

140 I18071, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 2000-1000 BCE, HRV_Bezdanjača_BA, J1c2, R1b1a1b-M269

141 I18086, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 2000-1000 BCE, HRV_Bezdanjača_BA, n/a 

142 I18075, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 2000-1000 BCE, HRV_Bezdanjača_BA, T2c1a2, R1b-L754

143 I5074, Lazaridis et al. 2022, Dalmatia, Matkovici, Middle-Late Bronze Age, 1497-1397 BCE, HRV_BA, X2m/X2m3, J2b2a1a1a1b-Z38240

144 I18736, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_MLBA, n/a 

145 I18735, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_Bezdanjača_BA, U5a1a1 

146 I18734, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_Bezdanjača_BA, T2a1b1 

147 I18730, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_Bezdanjača_BA, U5b2b5 

148 I18729, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_Bezdanjača_BA, T2b, R1b1a1b1a1a2b1c1a1a2a-BY150850

149 I18724, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_Bezdanjača_BA, HV0e 

150 I18732, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_Bezdanjača_BA, HV24, R1b1a1b1a1a2b1-L2

151 I18727, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_Bezdanjača_BA, T2c1d 

152 I18725, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_Bezdanjača_BA, V10b1, R1b1a1b1a1a2b1-L2

153 I18738, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_Bezdanjača_BA, H5a6 

154 I18413, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_Bezdanjača_BA, H1 

155 I18718, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_MLBA, n/a 

156 I18739, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_Bezdanjača_BA, T2c1d 

157 I18728, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_Bezdanjača_BA, T2c1d

158 I18414, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_Bezdanjača_BA, n/a 

159 I18717, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_Bezdanjača_BA, H5r, R1b1a1b1a1a2b1-L2

160 I18417, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_Bezdanjača_BA, X2b, R1b1a1b1a1a2b1-L2

161 I18733, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_Bezdanjača_BA, H1at 

162 I18720, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_Bezdanjača_BA, U8b1b1 

163 I18722, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_Bezdanjača_BA, U5a 

164 I18415, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_Bezdanjača_BA, H1aj1, R1b1a1b1a1a2b1-L2

165 I18737, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_Bezdanjača_BA, U5b1c1a, R1b1a1b1a1a2b1-L2

166 I18416, Lazaridis et al. 2022, Lika, Bezdanjača Cave, Middle-Late Bronze Age, 1500-800 BCE, HRV_Bezdanjača_BA, H3 

167 E3919, Marjanović et al. 2026, Dalmatia, Krneza-Duševića Glavica, Early Bronze Age, I1a1 

168 E3922, Marjanović et al. 2026, Dračevac Tumul, Early-Middle Bronze Age, T2a1b, J2b-Y31143

169 E3916, Marjanović et al. 2026, Ljubač Nekropola, Bronze Age, K1ag, J2b-Z2432

170 E3921, Marjanović et al. 2026, Vrbica Tumul, Bronze Age, J2b1C

171 E3920, Marjanović et al. 2026, Vrbica Tumul, Bronze Age, K1a

172 E3925, Marjanović et al. 2026, Torine Tumul, Bronze Age, U3a1, J2a-Y20492

173 E3926, Marjanović et al. 2026, Torine Tumul, Bronze Age, U4a2, A1*

174 E3914, Marjanović et al. 2026, Ljubač Nekropola, Late Bronze / Early Iron Age, HV0e

175 E3912, Marjanović et al. 2026, Ljubač Nekropola, Late Bronze / Early Iron Age, HV0e

176 I24344, Patterson et al. 2022, Dalmatia, Velim-Kosa, Middle-Late Bronze Age / Early Iron Age, 1500-400 BCE, Croatia_MBA_LBA_EIA, H13a2a 

177 I24345, Patterson et al. 2022, Dalmatia, Velim-Kosa, Middle-Late Bronze Age / Early Iron Age, 1500-400 BCE, Croatia_MBA_LBA_EIA, HV0e, J2b2a1-L283

178 I3313, Mathieson et al. 2018/Patterson at al. 2022, Dalmatia, Jazinka Cave, Late Bronze Age-Early Iron Age, 805-761 BCE, Croatia_LBA, HV0e

179 E3910, Marjanović et al. 2026, Ljubač Nekropola, Early Iron Age, H2, R1b-Z2108 (pogrešno upisan R1a)

180 E3911, Marjanović et al. 2026, Ljubač Nekropola, Early Iron Age, H2a2, R1b-L23

181 E3915, Marjanović et al. 2026, Ljubač Nekropola, Early Iron Age, HV0e

182 E3908, Marjanović et al. 2026, Ljubač Nekropola, Early Iron Age, N1b1a

183 E3907, Marjanović et al. 2026, Ljubač Nekropola, Early Iron Age, U2d2, J2b-Z1825 (pogrešno upisan J1a)

184 E3913, Marjanović et al. 2026, Ljubač Nekropola, Early Iron Age, H2a 

185 E3909, Marjanović et al. 2026, Ljubač Nekropola, Early Iron Age, n/a

186 E3924, Marjanović et al. 2026, Torine Tumul, Early Iron Age, H4d

187 E3923, Marjanović et al. 2026, Torine Tumul, Early Iron Age, T2b, J2b-YP50

188 I5729, Lazaridis et al. 2022, Continental, Sv. Petar Ludbreški, Early Iron Age, 1051-901 BCE, HRV_IA M, V, G2a2b2a1a1b-L497

189 I23911, Patterson et al. 2022, Lika, Smiljan, Early Iron Age, 891-797 BCE, Croatia_EIA, T2a1a, J2b2a1a1a1b2-Y86930

190 I23996, Patterson et al. 2022, Lika, Smiljan, Early Iron Age, 809-779 BCE, Croatia_EIA, H3b 

191 I23995, Patterson et al. 2022, Lika, Smiljan, Early Iron Age, 810-675 BCE, Croatia_EIA, H3b, J2b2a1a1a1b2 -Y86930

192 I24638, Patterson et al. 2022, Lika, Smiljan, Early Iron Age, 795-567 BCE, Croatia_EIA, U5a1g, J2b2a1a1a1b2 -Y86930

193 I24639, Patterson et al. 2022, Lika, Smiljan, Early Iron Age, 795-567 BCE, Croatia_EIA, H1, J2b2a1a1a1b2-Y86930

194 I23904, Patterson et al. 2022, Lika, Smiljan, Early Iron Age, 789-568 BCE, Croatia_EIA, T2b

195 I24883, Patterson et al. 2022, Kvarner, Osor-St. Peter, Early Iron Age, 850-750 BCE, Croatia_EIA, HV0e 

196 I26742, Patterson et al. 2022, Dalmatia, Jazinka Cave, Early Iron Age, 800-600 BCE, Croatia_EIA, HV0e, J2b2a1a1a1b2-PH1602

197 I24882, Patterson et al. 2022, Dalmatia, Mala Metaljka, Early Iron Age, 774-549 BCE, Croatia_EIA, H, J2b2a1a1a1b2-Y86930

198 I24879, Patterson et al. 2022, Dalmatia, Skradnik-Sultanov grob, Early Iron Age, 774-549 BCE, Croatia_EIA, H5/H5a1 

199 I5727, Lazaridis et al. 2022, Continental, Sv. Križ Brdovečki, Iron Age, 765-489 BCE, HRV_IA, J1c2

200 I5725, Lazaridis et al. 2022, Continental, Sv. Križ Brdovečki, Iron Age, 752-417 BCE, HRV_IA, H, R1b1a1b1a1a2b1c2-Y4355

201 I5726, Lazaridis et al. 2022, Continental, Sv. Križ Brdovečki, Iron Age, 538-399 BCE, HRV_IA, T1a2a 

202 I5723, Lazaridis et al. 2022, Continental, Sv. Križ Brdovečki, Iron Age, 514-391 BCE, HRV_IA, HV1a'b'c, J2b2a1-Z622

203 I5724, Lazaridis et al. 2022, Continental, Sv. Križ Brdovečki, Iron Age, 382-206 BCE, HRV_IA, U5b1d1a, E1b1b1a1b1a14-Y16721

204 I5728, Lazaridis et al. 2022, Continental, Sv. Križ Brdovečki, Iron Age, 361-200 BCE, HRV_IA, T1a5 

205 I18831, Lazaridis et al. 2022, Dalmatia, Velim-Kosa, Late Iron Age – Antiquity, 500-1 BCE, HRV_Anc, H7, J2b2a1a1a1b2-PH1602

206 I18830, Lazaridis et al. 2022, Dalmatia, Velim-Kosa, Late Iron Age – Antiquity, 500-1 BCE, HRV_Anc, H7, J2b2a1a1a1b2-PH1602

207 I26735, Patterson et al. 2022, Continental, Osijek (Vojarna), Late Iron Age, 150-50 BCE, Croatia_EIA_LaTene, J1c12, R1b-V88

208 I26706, Olalde et al. 2023, Dalmatia, Dragulin necropolis-Trogir, Roman period, 1-50 CE, n/a

209 I26702, Olalde et al. 2023, Dalmatia, Dragulin necropolis-Trogir, Roman period, 1-200 CE, H, E1b1b1a1b1a14-Y16729

210 I26708, Olalde et al. 2023, Dalmatia, Dragulin necropolis-Trogir, Roman period, 1-200 CE, H

211 I26703, Olalde et al. 2023, Dalmatia, Dragulin necropolis-Trogir, Roman period, 1-200 CE, T1a5a

212 I26704, Olalde et al. 2023, Dalmatia, Dragulin necropolis-Trogir, Roman period, 1-200 CE, T2b

213 I26705, Olalde et al. 2023, Dalmatia, Dragulin necropolis-Trogir, Roman period, 1-200 CE, C1

214 I26707, Olalde et al. 2023, Dalmatia, Dragulin necropolis-Trogir, Roman period, 1-200 CE 

215 HR3665, Antonio et al. 2024, Dalmatia, Dragulin necropolis-Trogir, Roman period, 124-217 CE, J2a1-S8230

216 R3670, Antonio et al. 2024, Dalmatia, Policija-Trogir, Roman period, 120-215 calCE, n/a 

217 I26775, Olalde et al. 2023, Dalmatia, Ulica Polačišće-Zadar, Roman period, 100-300 CE, H14a, J2a1a2b2 -Y13128

218 R3743, Antonio et al. 2024, Dalmatia, Ulica Polačišće-Zadar, Roman period, 81-210 CE, n/a, J2a-BY22478

219 R3744, Antonio et al. 2024, Dalmatia, Ulica Polačišće-Zadar, Roman period, 81-210 CE, n/a, R1b-P312

220 I26776, Olalde et al. 2023, Dalmatia, Poliklinika-Zadar, Roman period, 200-400 CE, H105, R1b1a1b1a1a2c1a-DF13

221 R3746, Antonio et al. 2024, Dalmatia, Poliklinika-Zadar, Roman period, 127-227 CE, n/a 

222 R3747, Antonio et al. 2024, Dalmatia, Poliklinika-Zadar, Roman period, 127-227 CE, n/a, R1b-Y3140

223 R3742, Antonio et al. 2024, Dalmatia, TC Relja-Zadar, Roman period, 127-227 CE, n/a, E-Y16161

224 R3745, Antonio et al. 2024, Dalmatia, Hypo Banka-Zadar, Roman period, 22-121 CE, n/a, E1b1b-BY66293

225 POP23, Freilich et al. 2021, Continental, Beli Manastir-Popova zemlja, Roman period, 260-402 CE, Croatia_Pop_RomanP, T2f2c, R1a1a1b2a2b1-F1019

226 R3542, Antonio et al. 2024, Continental, Beli Manastir, Roman period, 255-405 CE, n/a, R1b-Y39302

227 I26709, Olalde et al. 2023, Dalmatia, Dobrić necropolis-Trogir, Roman period-Early Middle Ages, 300-600 CE, H15a1 

228 I26714, Olalde et al. 2023, Dalmatia, Dobrić necropolis-Trogir, Roman period, 300-400 CE, K1a

229 SK 1, Novak et al. 2025, Continental, Mursa-Osijek, Roman period, n/a 

230 SK 2, Novak et al. 2025, Continental, Mursa-Osijek, Roman period,  232-335 CE, T2b5, N1a1a1a1a1a1a 

231 SK 3, Novak et al. 2025, Continental, Mursa-Osijek, Roman period, K1a3a1, R1b1a1b1a1a1c1a

232 SK 4, Novak et al. 2025, Continental, Mursa-Osijek, Roman period, 244-375 CE, T2b, I2a2a-M223

233 SK 5, Novak et al. 2025, Continental, Mursa-Osijek, Roman period, 239-346 CE, n/a

234 SK 6, Novak et al. 2025, Continental, Mursa-Osijek, Roman period, n/a

235 SK 7, Novak et al. 2025, Continental, Mursa-Osijek, Roman period, 240-340 CE, K1a4c, I1a3a1

236 I26718, Olalde et al. 2023, Dalmatia, Dobrić necropolis-Trogir, Roman period, 100-400 CE, V16, R1b1a1b1b3a1a1b-BY250

237 I26719, Olalde et al. 2023, Dalmatia, Dobrić necropolis-Trogir, Roman period, 100-400 CE, H27a

238 I26717, Olalde et al. 2023, Dalmatia, Dobrić necropolis-Trogir, Roman period-Early Middle Ages, 300-600 CE, c/a 

239 I26713, Olalde et al. 2023, Dalmatia, Dobrić necropolis-Trogir, Roman period, 100-500 CE, c/a

240 I26711, Olalde et al. 2023, Dalmatia, Dobrić necropolis-Trogir, Roman period-Early Middle Age, 300-600 CE, n/a

241 I26750, Olalde et al. 2023, Continental, Ban Jelačić Square-Osijek, Roman period, 200-300 CE, H5d 

242 I26751, Olalde et al. 2023, Continental, Ban Jelačić Square-Osijek, Roman period, 200-300 CE, H79 

243 I26752, Olalde et al. 2023, Continental, Ban Jelačić Square-Osijek, Roman period, 200-300 CE, K1b1a1a, J2b2a1a1a1a1a2a-Y23094

244 I26753, Olalde et al. 2023, Continental, Ban Jelačić Square-Osijek, Roman period, 200-300 CE, H, G2a2b2a1a1a1a-L13

245 I26754, Olalde et al. 2023, Continental, Ban Jelačić Square-Osijek, Roman period, 200-300 CE, H41 

246 R3655, Antonio et al. 2024, Continental, Ban Jelačić Square-Osijek, Roman period, 133-306 CE, G2a-S18765

247 R3656, Antonio et al. 2024, Continental, Ban Jelačić Square-Osijek, Roman period, 229-329 CE, n/a

248 R3657, Antonio et al. 2024, Continental, Ban Jelačić Square-Osijek, Roman period, 229-329 CE, n/a

249 BBC001, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, H33c

250 BBC002, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, H76

251 BBC003, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, H33c

252 BBC004, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, I5c, G2a2b2a1a1b1-CTS9737/Z1815

253 BBC005, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, H4a1b1, J1a2a1a2d2b-CTS5269/PF4855/Z2322

254 BBC006, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, HV0e

255 BBC007, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, W1i1a

256 BBC008, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, H7c1, J2b2a1a1a-AM00216/AMM125/Z615

257 BBC009, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, H104a/H1-f2, J2b2a1a1a1b-CTS9215

258 BBC010, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, H33c

259 BBC011, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, H1-f2

260 BBC012, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, H13b1-a, J2b2a1a1a-AM00216/AMM125/Z615

261 BBC013, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, T1a-a

262 BBC014, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, H3/H76, G-CTS10945/M3206

263 BBC015, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, H5d, J2b-PF7336/Z540

264 BBC016, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, H33c

265 BBC017, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, K1a3a3, J2b2a1a1a1a1a-Z1297

266 BBC018, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, H7c1, J2b-PF7359/Z2419

267 BBC019, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, HV7, J2b2a1a1a1-Z2507

268 BBC020, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, H1-f2

269 BBC021, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, H1-f2, G2a-L31/PF3142/S149

270 BBC022, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, H3/H76, G2a2b-M3412/PF3278

271 BBC023, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, H5

272 BBC024, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, I5c, G2a-PF3141

273 BBC026, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, R1a1a

274 BBC027, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, K2b1b, J2b2a1a1a1b-Y15058/Z34462

275 BBC029, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, W1r, J2b2a-AM01378/Z625

276 BBC030, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, HV0e

277 BBC031, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, I5c, G2a-PF3141

278 BBC032, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, K1a3a3, J2b2a-AM01367/Z605

279 BBC033, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, I5c, G2a-PF3141

280 BBC034, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, I5c, G2a2b2a1a1b-S20969/Z3390

281 BBC035, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, J1c2

282 BBC037, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, H1-f2

283 BBC038, Gretzinger et al. 2025, Continental, Bubi's cave-Karlovac, Roman period, 250-300 CE, U8a1a1, E1b1b1a1b1-FGC11422/SK887/Y3865

284 I26761, Olalde et al. 2023, Continental, Pogorelec-Sisak, Roman period, 300-450 CE, T2b34

285 I26762, Olalde et al. 2023, Continental, Pogorelec-Sisak, Roman period, 300-450 CE, N1a2

286 I26763, Olalde et al. 2023, Continental, Pogorelec-Sisak, Roman period, 300-450 CE, H1e6, R1b1a1b1a1a2a2a-L617

287 I26764, Olalde et al. 2023, Continental, Pogorelec-Sisak, Roman period, 300-450 CE, U1a1a

288 I28446, Olalde et al. 2023, Continental, Pogorelec-Sisak, Roman period, 300-450 CE, H

289 I28447, Olalde et al. 2023, Continental, Pogorelec-Sisak, Roman period, 300-450 CE, T2c1d1

290 I28448, Olalde et al. 2023, Continental, Pogorelec-Sisak, Roman period, 300-450 CE, H13b1b

291 R2040, Antonio et al. 2024, Continental, Pogorelec-Sisak, Roman period, 245-402 CE, J2b-L283 >FT108131

292 R2041, Antonio et al. 2024, Continental, Pogorelec-Sisak, Roman period, 245-361 CE, J2a2-Z39727>FT151551

293 R2042, Antonio et al. 2024, Continental, Pogorelec-Sisak, Roman period, 252-402 CE, n/a

294 I35009, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, E1b1b1a1b1-L618

295 I34297, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, U5b1b, J2a1a1a2b2a3b1a1b1a2-Y23162

296 I34292, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, U5b1b

297 G13-fail, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, n/a

298 I34296, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, HV11, E1b1b1a1b1-L618

299 I34301, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, n/a

300 I34298, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, n/a

301 G17-fail, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, n/a

302 I34982, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, n/a

303 I34981, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, n/a

304 I34299, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, n/a

305 I34294, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, W3a1, J2b2a1a1a1a1a1a-Z631

306 I33811, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, U5b1b

307 G6-fail, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, n/a

308 I33812, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, U1a3

309 I35082, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, n/a

310 I34287, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, n/a

311 I33889, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, H35, J1a2a1a1-ZS4393

312 G12-A10-fail, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, n/a 

313 I34291, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, n/a

314 I34290, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, U8b1a1 

315 I33885, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, H3k

316 I33808, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, H3k, J2a1a1b2a1b1b2c-PH185

317 G12-A4-fail, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, n/a 

318 G12-A5-fail, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, n/a 

319 I33888, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, H9a, J2a1a4b-L25

320 I33886, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, HV, E1b1a1a1a1c1b2a2-Z22293

321 G12-A9-fail, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, n/a 

322 I33809, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, HV12b1, J2a1a1a2b1b-M319

323 I33893, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, H5a3a2 

324 I33890, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, R0a, G2a2b2a1a1b2-Z27264

325 I33891, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, K1c1

326 G12-A8-fail, Zagorc et al. 2024, Dalmatia, Radošević-Hvar, Roman period, 3rd-5th century CE, n/a 

327 I26896, Olalde et al. 2023, Dalmatia, Gardun, Roman period-Early Middle Ages, 400-600 CE, n/a

328 I26894, Olalde et al. 2023, Dalmatia, Gardun, Roman period-Early Middle Ages, 360-600 CE, H60a 

329 I26895, Olalde et al. 2023, Dalmatia, Gardun, Roman period-Early Middle Ages, 360-600 CE, T2g1a 

330 I26897, Olalde et al. 2023, Dalmatia, Gardun, Roman period-Early Middle Ages, 400-600 CE, H1e, R1b1a1b1a1a2b1-L2

331 I26899, Olalde et al. 2023, Dalmatia, Gardun, Roman period-Early Middle Ages, 432-598 CE, E1b1b1a1b1-L618

332 R3543, Antonio et al. 2024, Dalmatia, Gardun, Roman period-Early Middle Ages, 431-600 CE, n/a

333 R3544, Antonio et al. 2024, Dalmatia, Gardun, Roman period-Early Middle Ages, 549-600 CE, n/a

334 R3545, Antonio et al. 2024, Dalmatia, Gardun, Roman period-Early Middle Ages, 431-542 CE, n/a

335 I24343, Olalde et al. 2023, Dalmatia, Mala Metaljka, Roman period-Early Middle Ages, 433-578 CE, J1c5 

336 R2050, Antonio et al. 2024, Dalmatia, Omišalj-Mirine, Roman period-Early Middle Ages, 402-533 CE, n/a 

337 R2051, Antonio et al. 2024, Dalmatia, Omišalj-Mirine, Roman period-Early Middle Ages, 402-533 CE, n/a 

338 R2053, Antonio et al. 2024, Dalmatia, Omišalj-Mirine, Roman period-Early Middle Ages, 402-533 CE, n/a 

339 R3659, Antonio et al. 2024, Continental, Šćitarjevo, Roman period, 248-378 CE, E1b1b-V13> >Z38334

340 R3660, Antonio et al. 2024, Continental, Šćitarjevo, Roman period-Early Middle Ages, 433-560 CE, n/a, I1-Y151828

341 R3685, Antonio et al. 2024, Dalmatia, Velić, Roman period-Early Middle Ages, 434-565 CE, G2a- L497>FGC8346

342 SU 259, Fernandes et al. 2019, Continental, Hermanov vinograd-Osijek, Roman period-Early Middle Ages, 5th century CE, n/a 

343 SU 261, Fernandes et al. 2020, Continental, Hermanov vinograd-Osijek, Roman period-Early Middle Ages, 5th century CE, n/a 

344 SU 750, Fernandes et al. 2021, Continental, Hermanov vinograd-Osijek, Roman period-Early Middle Ages, 5th century CE, n/a 

345 E3917, Marjanović et al. 2026, Dalmatia, Krneza Duševića glavica, Early Middle Ages, H5a1, R1a-M459*

346 E3918, Marjanović et al. 2026, Dalmatia, Krneza Duševića glavica, Early Middle Ages, H5a1

347 R3547, Antonio et al. 2024, Dalmatia, Novo selo-Bunje, Early Middle Ages, 545-597 CE, n/a

348 I26765, Olalde et al. 2023, Istria, Sipar-Umag, Early Middle Ages, 650-800 CE, U8b1b1, G2a2b2a4a1a1b-Y7538

349 I26766, Olalde et al. 2023, Istria, Sipar-Umag, Early Middle Ages, 650-800 CE, HV1a'b'c4*, R1a1a1b1a2b3a3a2g2b1-YP4966

350 I26767, Olalde et al. 2023, Istria, Sipar-Umag, Early Middle Ages, 500-700 CE, K1a19, G2a2a1a2a1a1-Z6494

351 R2045, Antonio et al. 2024, Istria, Sipar-Umag, Early Middle Ages, 550-603 CE, n/a 

352 R3662, Antonio et al. 2024, Istria, Sipar-Umag, Early Middle Ages, 558-639 CE, n/a

353 R3663, Antonio et al. 2024, Istria, Sipar-Umag, Early Middle Ages, 686-876 CE, n/a

354 R3664, Antonio et al. 2024, Istria, Sipar-Umag, Early Middle Ages, 679-823 CE, E1b1b-V13>BY152493 www.yfull.com/tree/E-FT1910/

355 I26745, Olalde et al. 2023, Continental, Jagodnjak–Krčevine–Selska Bara-Beli Manastir, Early Middle Ages, 670-775 CE, J1c2 

356 I26746, Olalde et al. 2023, Continental, Jagodnjak–Krčevine–Selska Bara-Beli Manastir, Early Middle Ages, 600-850 CE, H9a, G2a2a1a2a1a1-Z6494

357 I26747, Olalde et al. 2023, Continental, Jagodnjak–Krčevine–Selska Bara-Beli Manastir, Early Middle Ages, 671-774 CE, H1

358 I26748, Olalde et al. 2023, Continental, Brekinjova kosa-Bojna-Glina, Early Middle Ages, 773-890 CE, I4a1, R1a1a1b1a1a1a-YP1337

359 I26749, Olalde et al. 2023, Continental, Brekinjova kosa-Bojna-Glina, Early Middle Ages, 774-885 cal CE, W3b, J1a2a1a2d2b2b2c4d-Y4349

360 I28388, Olalde et al. 2023, Continental, Dvorac-Nuštar, Early Middle Ages, 700-900 CE, U8b1a1, E1b1b1a1b1a10b-FGC11457

361 I28390, Olalde et al. 2023, Continental, Dvorac-Nuštar, Early Middle Ages, 771-947 CE, H1 

362 I28392, Olalde et al. 2023, Continental, Dvorac-Nuštar, Early Middle Ages, 700-900 CE, H7a1a

363 I28393, Olalde et al. 2023, Continental, Dvorac-Nuštar, Early Middle Ages, 700-900 CE, J1c2

364 I28394, Olalde et al. 2023, Continental, Dvorac-Nuštar, Early Middle Ages, 700-900 CE, H5 

365 I28397, Olalde et al. 2023, Continental, Dvorac-Nuštar, Early Middle Ages, 700-900 CE, J1c1b1a, J2b2a1a1a1a1a1a1-Z1043

366 I28398, Olalde et al. 2023, Continental, Dvorac-Nuštar, Early Middle Ages, 700-900 CE, K1a4j

367 I28400, Olalde et al. 2023, Continental, Dvorac-Nuštar, Early Middle Ages, 700-900 CE, H14b2, Q1a2a1-YP844

368 I34800, Olalde et al. 2023, Continental, Dvorac-Nuštar, Early Middle Ages, 700-900 CE, H16, J2b2a1a1a1a1a1a-Z1043

369 I35081, Olalde et al. 2023, Continental, Čepinski Martinci, Early Middle Ages, 650-870 CE, U4a3a 

370 I15742, Lazaridis et al. 2022/Olalde et al. 2023, Dalmatia, Matije Gupca 15 street-Trogir, Early Middle Ages, 661-775 CE, HRV_Trogir_Byz_outlier, H3h6 

371 I15744, Lazaridis et al. 2022/Olalde et al. 2023, Dalmatia, Matije Gupca 15 street-Trogir, Early Middle Ages, 674-774 CE, HRV_Trogir_Byz, X2e2b 

372 I15462, Lazaridis et al. 2022/Olalde et al. 2023, Dalmatia, Matije Gupca 15 street-Trogir, Early Middle Ages, 677-824 CE, HRV_Trogir_Byz, T1a1 

373 I15463, Lazaridis et al. 2022/Olalde et al. 2023, Dalmatia, Matije Gupca 15 street-Trogir, Early Middle Ages, 700-800 CE, HRV_Trogir_Byz, X2e2b 

374 I15741, Lazaridis et al. 2022/Olalde et al. 2023, Dalmatia, Matije Gupca 15 street-Trogir, Early Middle Ages, 771-952 CE, HRV_Trogir_Byz, H 

375 I15743, Lazaridis et al. 2022/Olalde et al. 2023, Dalmatia, Matije Gupca 15 street-Trogir, Early Middle Ages, 774-979 CE, HRV_Trogir_Byz, H7d 

376 VEM001, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 700-900 CE, U5a1g, I2a1a2b1a1a-S20602/YP196

377 VEM002, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, H3-c

378 VEM003, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 702-879 CE, J1c5, N1a1a1a1a1a1a1-B215/L1025

379 VEM004, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, K1a4c

380 VEM005, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, K1a4c

381 VEM006, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 774-885 CE, H/H3-c, R1a1a1b1a2b-CTS1211/S3357

382 VEM007, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, H1a, I2a1a2b1a1a-S20602/YP196

383 VEM008, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, V7a1, I1a1b1a4a2a-Y15505

384 VEM009, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, H+152/H32, J2b2a1a1a1b-CTS9215

385 VEM010, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, U7b/U7b19

386 VEM011, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, U5a1b1c

387 VEM012, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, J1c2k

388 VEM013, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 682-876 CE, T2b5, R1a1a1b1a2b-CTS1211/S3357

389 VEM014, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 782-882 CE, R/U5

390 VEM015, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, T2b4e/R13, J2b2a1a1a-AM00216/AMM125/Z615

391 VEM016, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 688-877 CE, V4, I1c1c1a-A9726

392 VEM017, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 706-879 CE, U5b1

393 VEM018, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 646-672 CE, I5a, I2a1a2b1a1a-S20602/YP196

394 VEM019, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 772-890 CE, NA/N14

395 VEM020, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 773-883 CE, H46/H

396 VEM021, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 685 - 876 CE, V4, I1c1c1a-A9726

397 VEM022, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 775-884 CE, T2a1a1/T2a1a1b, E1b1b1a1b1a-L542/PF2220

398 VEM023, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650 CE - 900 CE, T2b5

399 VEM024, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650 CE - 900 CE, I/I3a, J2b2a1a1a1b-CTS9215

400 VEM025, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650 CE - 900 CE, I5a/I14, J2b2a1a1a1a1a1a1e-CTS11760

401 VEM026, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 710-883 CE, K1a4c1

402 VEM027, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 772 - 947 CE, U5b1e1, I2a1a2b1a1-FGC12096/Y3121

403 VEM028, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 707-881 CE, U2e1b1/U2e1b1a

404 VEM029, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 771-954 CE, H6a1a/H6a1a12, I2a1b2a2b2-BY14034/Y31042

405 VEM030, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, H5a1m

406 VEM031, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, H1f/H1f-a

407 VEM032, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 708-882 CE, H7f/H-d17, N1a1a1a1a1a1a1-B215/L1025

408 VEM033, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 773-980 CE, I5a/I5a1, J2b2a1a1a1a1a1a1e-CTS11760

409 VEM034, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, M31a2b2 

410 VEM035, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 892-992 CE, H36/H36a, R1a1a1b1a2b3a-CTS3402/S3361/V2670

411 VEM036, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, H6a1a

412 VEM037, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, H1c

413 VEM038, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, V6, R1b1a1b1b3a1a1a-Y5587

414 VEM039, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 774 - 885 CE, U5b1

415 VEM040, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 772-959 CE, H6a1a/H6a1a4b3

416 VEM041, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, H84/H84a

417 VEM042, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, J2b1c

418 VEM043, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 772-884 CE, U4d1a1a1, I2a1a2b1a1-FGC12076/YP217

419 VEM044, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 707-880 CE, J1c5/J1c

420 VEM045, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, B5a1c1a4

421 VEM046, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, n/a

422 VEM047, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, H5a2/H5a

423 VEM048, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, K1/K1-a, J2b2a1a1a1b-Y15058/Z34462

424 VEM049, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 687-878 CE, H5a2, R1a1a1b1a2b3a-FGC4501/V2834/Y2194

425 VEM050, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 773-883 CE, H2/H2a2a1i

426 VEM051, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, K2a/U8b, J2b2a1a1a1b-CTS9215

427 VEM052, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, W1e1a/W1e1a1, I2a1a2b1a1a-S20602/YP196

428 VEM053, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 773-883 CE, H1c22

429 VEM054, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 673-773 CE, K1a4a1b/K1a4a1f, R1a1a1b1a1a1a-L260/S222

430 VEM055, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, J1c3/J1c3-a1

431 VEM056, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 704-879 CE, U5b1b1a, I1a1b1a1b3a-FGC21579

432 VEM057, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, U5b1b1a, I1a1b-CTS10028/Z2337

433 VEM058, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 705-879 CE, I1a1b1a1b3a-FGC21579

434 VEM059, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, J1c5/J1c2u1a, I1a-CTS9857/V3772

435 VEM060, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 774-884 CE, H13a1c

436 VEM061, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 673-775 CE, HV2a1/HV2a1a

437 VEM062, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, K1a2

438 VEM063, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, U4c

439 VEM064, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, H5a2/H5a2f, J2b2a1a1a1b-Y15058/Z34462

440 VEM065, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, P5/R0a1b

441 VEM066, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, T2b5

442 VEM067, Gretzinger et al. 2025, Dalmatia, Velim, Early Middle Ages, 650-900 CE, J1b1a1b

443 POD001, Gretzinger et al. 2025, Dalmatia, Podvršje-Glavčine, Early Middle Ages, 400-850 CE, L3e5d

444 POD002, Gretzinger et al. 2025, Dalmatia, Podvršje-Glavčine, Early Middle Ages, 400-850 CE, H13a1a1/H1dd

445 VIR001, Gretzinger et al. 2025, Dalmatia, Vir-Smratina, Early Middle Ages, 400-1200 CE, H32b1

446 DUC001, Gretzinger et al. 2025, Dalmatia, Dubrovnik cathedral, Early Middle Ages, 500-1100 CE, U5a1m1, I2a1a2b1a1-S15187/YP191

447 BIO001, Gretzinger et al. 2025, Dalmatia, Biograd-Glavice, Middle Ages, 1200-1700 CE, W4a1

448 BRI006, Gretzinger et al. 2025, Dalmatia, Bribir-Glavica, Middle Ages, 1200-1700 CE, H5

449 DUG014, Gretzinger et al. 2025, Dalmatia, Dugopolje-Vučipolje, Middle Ages, 1200-1600 CE, K1-a17, I2a1a2b1a1-CTS5996

450 TOR001, Gretzinger et al. 2025, Continental, Torčec-Cirkvišće, Middle Ages, 1100-1800 CE, J2b1-J2b1k

451 TOR002, Gretzinger et al. 2025, Continental, Torčec-Cirkvišće, Middle Ages, 1100-1800 CE, V18a2

452 TOR003, Gretzinger et al. 2025, Continental, Torčec-Cirkvišće, Middle Ages, 1100-1800 CE, H5a, I2a1a2b-S26147

453 TOR004, Gretzinger et al. 2025, Continental, Torčec-Cirkvišće, Middle Ages, 1100-1800 CE, I2/I2d1b1f, I2a1a2b1a1a1-Y3548

454 TOR005, Gretzinger et al. 2025, Continental, Torčec-Cirkvišće, Middle Ages, 1100-1800 CE, H5a1

455 TOR006, Gretzinger et al. 2025, Continental, Torčec-Cirkvišće, Middle Ages, 1100-1800 CE, T2b14a 

456 I35012, Olalde et al. 2023, Lika, Gornji Kosinj-Saint Ana, Middle Ages, 1161-1264 CE, U5b2a1a1, R1a1a1b1a2b-YP971

457 I34980, Olalde et al. 2023, Lika, Gornji Kosinj-Saint Ana, Middle Ages, 1100-1400 CE, U5b2a1a1

458 I35014, Olalde et al. 2023, Lika, Gornji Kosinj-Saint Ana, Middle Ages, 1000-1200 CE, T2h2, R1b1a1b1b3a1a1a-Y14300

459 I35095, Olalde et al. 2023, Lika, Gornji Kosinj-Saint Ana, Middle Ages, 1000-1200 CE, U3a1

460 I35010, Olalde et al. 2023, Lika, Gornji Kosinj-Saint Ana, Middle Ages, 1000-1200 CE, U5b1f

461 I1879, Lazaridis et al. 2022, Continental, Vinkovci-Jugobanka, Late Middle Ages-Early Modern Age, 1468-1636 CE, HRV_Vinkovci_Mdv, V6, I2a1a2b1a1-CTS10228

462 I35008, Olalde et al. 2023, Lika, Gornji Kosinj-Saint Ana, Early Modern Age, 1650-1750 CE, T1a1l, R1a1a1b1a2b3a4a2d-YP613

Some additional unpublished tested samples from 2024 are expected from early medieval Jagodnjak and Šarengrad (both predominantly E1b1b).

Misdated:

463 I18719, Patterson et al. 2022, Lika, Bezdanjača Cave, NOT Middle-Late Bronze Age (YES Middle Ages or Modern), NOT 1500-1000 BCE, NOT Croatia_MBA_LBA, HV0a1a/HV0a1a1b, I2a1a2b1a1-CTS10228>Y3120

464 I18721, Lazaridis et al. 2022, Lika, Bezdanjača Cave, NOT Middle-Late Bronze Age (YES Middle Ages or Modern), NOT 1500-800 BCE, NOT HRV_Bezdanjača_BA, T2b11, I2a1a2b1a1-Y3120>S17250>BY128/Y5596

465 I18723, Lazaridis et al. 2022, Lika, Bezdanjača Cave, NOT Middle-Late Bronze Age (YES Middle Ages or Modern), NOT 1500-800 BCE, NOT HRV_Bezdanjača_BA, V1a1, R1b1a1b1a1a2-P312